Cairngorms, Speyside Deer Management Group

Part 4 - Tactical and Operational Objectives, Cull Targets and Data Requirements

11.1 Requirements for mature stags (defined as stags of 6 years and older)
Trophy quality mature stags for sporting are usually over 6 years old. Clearly any specific requirement to provide these will require a higher population than if all ages of stags are acceptable in the cull. The following calculation is aimed at assessing the population required to provide the stag culls required by estates. These stag culls are based largely on previous culls, which were considered necessary to fulfil socio-economic objectives. This calculation illustrates any shortfall between requirements and availability, given the aims of reducing populations to achieve the primary objective. An aspiration by an estate to provide a particular number of mature or trophy quality stags does not influence the population models.

Stag requirements based on data provided by estates (Appendix 8) are presented below. For those estates not specifying a requirement for mature stags approximately one third of the total stag requirement has been assumed to allow calculation of the minimum population size;

Western sub-area    
  Stag cull required
  Total stags Mature stags
Total 325 170

Eastern sub-area    
  Stag cull required
Estate Total stags Mature stags
Total 95 33

The population required to support the required stag cull has been estimated as follows:
 
Western sub-area

170 x 7.6 = 1292 stags
1292 x 1.3 = 1680 hinds
1680 x 0.37 = 622 calves
Total population required = 3594

Eastern sub-area

33 x 7.6 = 251 stags
326 x 1.3 = 424 hinds
424 x 0.37 = 157 calves
Total population required = 832

 
NB This is based on life tables of red deer (Ratcliffe, 1987), which suggest that approximately 0.33 of the population will be aged 5 years and over. If we then assume that 0.4 of this part of the population will be actually available to client stalkers (ie it will be possible to contact these deer), 0.33 x 0.4 = 0.132 of the population are available trophy stags. Thus 0.132/1.0=7.6

NB Blank stalking days will be recorded by stalkers to monitor the actual availability of stags

Hinds are based on an assumed adult sex ration of 1.3 hinds: 1 stag, given the degree of under-culling and the higher survival rates of hinds.

Recruitment of calves of 0.37 is based on previous values for high performance red deer populations.

 
11.2 Current population size
Tables 3 and 4 are derived from initial information provided by estates, from subsequent discussions and from DCS February 2005 count data and dung counts for Forest Enterprise at Inshriach. Where no figure is available, low ground area has been estimated at 0.66 of total area.
 
Table 3 CSDMG western sub area plus Dalnacardoch, Athol and Mar Lodge – Summary of deer data
Estate Area
(total) (ha)
Area
(low)
(ha)
Stag
count
Unclassified
counted
Total stags (counted plus 0.05 of u/c) hinds (u/c – u/c stags / 1.35) calves (hinds x .35) Density
(total)
(Nkm-2)

Density
(low)
(Nkm-2)

Total CSDMG 64846 44811 1291 3921 5362 1487 2759 966 8.3 12.0
Total inc Athol, Dalnacardoch and Mar Lodge 123828 83739 2808 10185 13143 3318 7167 2508 10.6 15.7

Unclassified deer are divided on the basis that x 0.05 are juvenile stags (these are added to the counted stags) and the remainder are hinds and calves. Calves are assumed to be x 0.35 hinds. This follows DCS protocol for allocating unclassified deer from count data (I Hope, pers. comm.).

Thus the starting figures for the western sub-area (Table 5A) are 1487 stags and 2759 hinds and the totals including neighbouring estates (Table 5B) are 3318 stags and 7167 hinds.

 
Table 4 CSDMG eastern sub area plus Invercauld home beat - Summary of deer data
Estate Area
(total) (ha)
Area
(low)
(ha)
Stag
count
Unclassified
counted
Total stags (counted plus 0.05 of u/c) hinds (u/c – u/c stags / 1.35) calves (hinds x .35) Density
(total)
(Nkm-2)

Density
(low)
(Nkm-2)

Total CSDMG 39878 32997 564 1232 1118 422 506 190 2.8 3.4
Totals inc Invercauld 62328 47814 858 2404 2584 772 1331 479 4.1 5.4
 

Unclassified deer are divided on the basis that x 0.05 are juvenile stags (these are added to the counted stags) and the remainder are hinds and calves. Calves are assumed to be x 0.35 hinds. This follows DCS protocol for allocating unclassified deer from count data (I Hope, pers. comm.).

Thus the starting figures for the eastern sub-area (Table 7A) are 422 stags and 506 hinds. With the addition of the Invercauld home beat (Table 7B), the starting figures are 772 stags and 1331 hinds.

 
Cull targets and population models
Western sub-area
The stag requirements of estates in the western sub-area of 170 mature stags require a spring population of 3,594 red deer (equivalent density of 5.5 deer km-2). The DCS count in spring 2005, plus estimates from dung counts in Inshriach woodlands, etc., estimated 5362 deer (Table 3) (equivalent spring density of 8.3 km-2). This suggests that the current population can easily provide for the mature stags required by estates. Planned population reductions will close the gap between the actual population and that required to provide the mature stag culls but the reduced numbers indicated in 2010 (Table 5A) of 1487 stags, 1909 hinds and 706 calves still exceed those required to provide the sporting stag culls of estates.

However, it seems clear that deer move in and out of the area in the south and south-east and that an influx of stags, particularly to the Gaick and Glenfeshie hind populations, is helping to sustain stag requirements in the sub-area. In order to meet the CSDMG objectives (Section 10), it is proposed that the spring population should be around 9 deer km-2. This will allow all objectives to be met.

Three models are provided. Table 5A includes only those estates that are part of the CSDMG area and indicates a reduction in the spring density from 11.8 deer km-2 to 9.2 deer km-2. The starting population values in Tables 5 A-C, are the values of stags and hinds from Table 3 and a revised value for calves based on a recruitment rate of hinds of x 0.37 (this is the reason for the small discrepancy between the total population value in Figures 5 A-C and 7 A-B).

The concentrations occurring in the lower glens in periods of adverse weather may continue to cause unacceptable impacts on woodland regeneration and other vulnerable habitats and culling should aim to achieve localised differences in density relevant to local objectives. An important consideration is the increase in population size due to incursions from estates outside the CSDMG, notably from parts of Athol, Dalnacardoch and Mar Lodge estates. Because of the current imbalance in favour of hinds and in order to maintain stag numbers, stag culls are equivalent to recruitment, while hinds are culled more heavily. This results in an increase in the ratio of stags:hinds from 0.54 to 0.78. However, with the influx of stags from the south, summer sex ratios will be closer to parity.

 
Table 5A Red deer culling model for the CSDMG western sub-area
 
Year Population Parameter Stag Hind Calf Total Density
(km-2)*
Density
(km-2)**
2005 Spring count 1487 2759 1021 *5267   11.8
  Recruitment (Yearlings) 511 510        
  Summer (after calving) 1998 3269 1308 6575 10.1  
  Cull (S=recruitment; H=recruitment+150) 511 660 264 1435    
2006 Spring estimate 1487 2609 965 5061   11.3
  Recruitment (Yearlings) 483 482        
  Summer (after calving) 1970 3091 1236 6297 9.7  
  Cull (S=r; H=r+200) 483 682 273 1438    
2007 Spring estimate 1487 2409 891 4787   10.7
  Recruitment (Yearlings) 445 446        
  Summer (after calving) 1932 2855 1142 5929 9.1  
  Cull (S=r;H=r+200) 445 646 258 1349    
2008 Spring estimate 1487 2209 817 4516   10.1
  Recruitment (Yearlings) 408 409        
  Summer (after calving) 1895 2618 1047 5560 8.6  
  Cull (S=r;H=r+200) 408 609 244 1261    
2009 Spring estimate 1487 2009 743 4239   9.5
  Recruitment (Yearlings) 371 372        
  Summer (after calving) 1858 2381 952 5191 8.0  
  Cull (S=r;H=100) 371 472 189 1032    
2010 Spring estimate 1487 1909 706 4102   9.2
  Ongoing maintenance cull = recruitment 371 472 189 1032    

* Summer densities based on the entire available range of 64,846 ha.
** Spring densities are computed based on a winter range of 44,811 ha.

Assumes 0.4 of hinds in summer calving; 0.37 of calves recruited into adult population in spring..
Figures for the numbers of calves in the cull and in summer are crude estimates (hinds x.4) based on numbers of hinds. Calves should be culled in relation to milk hinds in the cull.
* The starting population values in Tables 5 A-C, are the values of stags and hinds from Table 3 and a revised value for calves based on a recruitment rate of hinds of x 0.37 (this is the reason for the small discrepancy between the total population value in Figures 5 and 7 A-C.

Table 5B presents a model including the neighbouring estates and aims to reduce the density from 15.7 deer km-2 to around 13 deer km-2. This final density assumes that increased culling (ie additional hind culls above recruitment; see Table 5B) will be focussed mainly in the CSDMG area and that estates outside the CSDMG area will be less prepared to reduce numbers to the same degree. Culls on estates within the CSDMG should approximately follow those proposed in Table 5A.

This is considered to be the more realistic model. However, its application will require acceptance and co-operation from the neighbouring estates. Although the CSDMG western sub area is subject to immigration from the neighbouring estates and this increases their availability of stags, any reductions outside the sub-area will result in an overall reduction in the availability of stags within it. The aim here (Table 5B) has been to maintain stag numbers, focussing culling pressure on hinds.

 
Table 5B Red deer culling model for the CSDMG western sub-area including Atholl, Dalnacardoch and Mar Lodge
Year Population Parameter Stag Hind Calf Total Density
(km-2)*
Density
(km-2)**
2005 Spring count 3318 7167 2652 13137*   15.7
  Recruitment (Yearlings) 1326 1326        
  Summer (after calving) 4644 8493 3397 16534 13.4  
  Cull (S=recruitment; H=recruitment+200) 1326 1526 610 3462    
2006 Spring estimate 3318 6967 2578 12863   15.4
  Recruitment (Yearlings) 1289

1289

       
  Summer (after calving) 4607 8256 3302 16165 13.1  
  Cull (S=r;H=r+200) 1289 1489 596 3374    
2007 Spring estimate 3318 6767 2504 12589   15.0
  Recruitment (Yearlings) 1252 1252        
  Summer (after calving) 4570 8019 3208 15797 12.8  
  Cull (S=r;H=r+200) 1252 1452 581 3285    
2008 Spring estimate 3318 6567 2430 12315   14.7
  Recruitment (Yearlings) 1215 1215        
  Summer (after calving) 4533 7782 3113 15428 12.5  
  Cull (S=r;H=r+200) 1215 1415 566 3196    
2009 Spring estimate 3318 6367 2356 12041   14.4
  Recruitment (Yearlings) 1178 1178        
  Summer (after calving) 4496 7545 3018 15059 12.2  
  Cull (S=r;H=r+200) 1178 1378 551 3107    
2010 Spring estimate 3318 6167 2282 11767   14.1
  Recruitment (Yearlings) 1141 1141        
  Summer (after calving) 4459 7308 2923 14690 11.9  
  Cull (S=r;H=r+200) 1141 1341 536 3018    
2011 Spring estimate 3318 5967 2208 11493   13.7
  Ongoing maintenance cull = recruitment 1104 1104 442 2650    
* Summer densities based on the entire available range of 123,828 ha.
** Spring densities are computed based on a winter range of 83,739 ha.
Assumes 0.4 of hinds in summer calving; 0.37 of calves recruited into adult population in spring.
Figures for the numbers of calves in the cull and in summer are crude estimates (hinds x 0.4) based on numbers of hinds. Calves should be culled in relation to milk hinds in the cull.

* The starting population values in Tables 5 A-C, are the values of stags and hinds from Table 3 and a revised value for calves based on a recruitment rate of hinds of x 0.37 (this is the reason for the small discrepancy between the total population value in Figures 5 and 7 A-C.

Table 5C presents a model for the wider area (as in Table 5B) but reduces the density to a similar level as presented for the CSDMG area alone (Table 5A). Again, culls on estates within the CSDMG should approximately follow those proposed in Table 5A. This model (Table 5C) will almost certainly fulfil CSDMG objectives but may not be acceptable to the neighbouring estates

 
Table 5C Red deer culling model for the CSDMG western sub-area including Atholl, Dalnacardoch and Mar Lodge aimed at reducing numbers to achieve a density similar to the CSDMG western sub area only (8.9 km-2; Table 5A) and restoring stag:hind ratio
Year Population Parameter Stag Hind Calf Total Density
(km-2)*
Density
(km-2)**
2005 Spring count 3318 7167 2652 13137*   15.7
  Recruitment (Yearlings) 1326 1326        
  Summer (after calving) 4644 8493 3397 16534 13.4  
  Cull (S=recruitment; H=recruitment+400) 1276 1726 690 3692    
2006 Spring estimate 3368 6767 2504 12639   15.1
  Recruitment (Yearlings) 1252 1252        
  Summer (after calving) 4620 8019 3208 15847 12.8  
  Cull (S=r;H=r+400) 1202 1652 661 3515    
2007 Spring estimate 3418 6367 2356 12141   14.5
  Recruitment (Yearlings) 1178 1178        
  Summer (after calving) 4596 7545 3018 15159 12.2  
  Cull (S=r;H=r+500) 1178 1678 671 3527    
2008 Spring estimate 3418 5867 2171 11456   13.7
  Recruitment (Yearlings) 1085 1086        
  Summer (after calving) 4504 6953 2781 14238 11.5  
  Cull (S=r;H=r+600) 1085 1686 674 3445    
2009 Spring estimate 3419 5267 1949 10635   12.7
  Recruitment (Yearlings) 974 975        
  Summer (after calving) 4394 6242 2497 1311 10.6  
  Cull (S=r;H=r+700) 974 1675 670 3319    
2010 Spring estimate 3420 4567 1690 9677   11.6
  Recruitment (Yearlings) 845 845        
  Summer (after calving) 4265 5412 2165 11842 9.6  
  Cull (S=r;H=r+700) 845 1545 618 3008    
2011 Spring estimate 3420 3867 1431 8718   10.4
  Recruitment (Yearlings) 715 716        
  Summer (after calving) 4135 4583 1833 10551 8.5  
  Cull (S=r;H=r+700) 715 1416 566 2697    
2012 Spring estimate 3420 3167 1172 7759   9.3
* Summer densities based on the entire available range of 123,828 ha.
** Spring densities are computed based on a winter range of 83,739 ha.

Assumes 0.4 of hinds in summer calving; 0.37 of calves recruited into adult population in spring.
Figures for the numbers of calves in the cull and in summer are crude estimates (hinds x 0.4) based on numbers of hinds. Calves should be culled in relation to milk hinds in the cull.

* The starting population values in Tables 5 A-C, are the values of stags and hinds from Table 3 and a revised value for calves based on a recruitment rate of hinds of x 0.37 (this is the reason for the small discrepancy between the total population value in Figures 5 and 7 A-C.

The model that includes the estates to the south of the western sub-group area but achieves an overall density of 13.7 deer km-2 (Model 5B) is considered to provide a more realistic vision of the red deer population. However, within the CSDMG area the culls proposed in Table 5A need to be achieved. It will be easier (but not impossible) to achieve the overall densities with agreement and co-operation from neighbouring estates and the CSDMG area will continue to import stags from the south and west. The estates in the south and west of the CSDMG area (notably, South Drumochter, Gaick and Glenfeshie) are dependent upon stags moving in from the south during the summer and reductions in deer numbers in Atholl, Dalnacardoch and Mar Lodge will reduce stag availability in the CSDMG area.

Tables 5A, 5B and 5C indicate a reduction in the total density in the western sub area aimed at maintaining stag numbers and reducing current impacts. Deer control should be concentrated on the sensitive areas (See 10 above) aimed at achieving local differences in density to deliver habitat-based targets in woodland regeneration areas and higher densities in largely sporting areas, whilst still permitting the recovery of important extensive habitats such as montane heath and tree-line scrub.

Local (estate-based) culls aimed at achieving the CSDMG western sub-area density of 9.2 deer km-2 and based on the initial years (2005-06) cull recommendation (line 8, Table 5A) are proposed in Table 6A. The division of the total cull requirement between estates has been based on the proportion of low ground (Table 3) as representing the approximate carrying capacity of the habitat. These proposed culls are indicative only and estates should share information on culling progress during culling seasons aimed at achieving the sub-area total.

 
Table 6A Indicative proposed red deer reduction culls for estates in the western sub-area
Estate % low ground Stag Hind Calf Total
(%)
Spring population
(from Table 3)
Total ** 100 483 682 273 1438 (27) 5362
*Estimated culls based on the proportion of low ground clearly do not reflect the dispersion of deer at particular times of year. Those marked with an * clearly do not currently carry sufficient deer in spring to justify the stated cull. However, the figures do indicate approximate cull requirements overall.

** Total cull figures are taken from line 8 of Table 5A (2006 culls)

Table 6B provides estimates of proposed sustainable culls following the reductions achieved (Table 6A) based on the earlier model (Table 5A).

 
Table 6B Proposed sustainable red deer culls for estates in the western sub-area following reduction to 4102 red deer (Table 5A)
Estate % low ground Stag Hind Calf Total
Total 100 371 472 189

1032

In practice cull figures should be shared between DMG members on a weekly basis during the cull season in order to achieve the overall sub-area targets.
Eastern sub-area

The stag requirements of estates in the eastern sub-area of 33 mature stags require a spring population of 832 red deer (equivalent spring density of 2.5 deer km-2). Starting values in Table 7A and 7B are derived from 2005 spring counts and recent reductions in Glenavaon. It appears that immigration from Invercauld is increasing stag availability during the stalking season. Given the low density in this area, the following models (Tables 7A and 7B) aim to maintain the total density in the eastern sub area as a means of maintaining sporting requirements. Deer control should be concentrated on the sensitive areas (See 10 above).

 
Table 7A Red deer culling model for the CSDMG eastern sub-area
Year Population Stag Hind Calf Total Density
(km-2)*
Density
(km-2)**
2005 Spring count 422 506 187 1115   3.4
  Recruitment (Yearlings) 93 94        
  Summer (after calving) 515 600 240 1355 3.4  
  Cull (=recruitment) 93 94 38 225    
2006 Spring estimate 422 506 187 1115   3.4
  Ongoing maintenance cull = recruitment 93 94 38 225    
* Summer densities based on the entire available range of 62,328 ha.
** Spring densities are computed based on a winter range of 47,814 ha.

Assumes 0.4 of hinds in summer calving; 0.37 of calves recruited into adult population in spring..
Figures for the numbers of calves in the cull and in summer are crude estimates based on numbers of hinds. Calves should be culled in relation to milk hinds in the cull.

 
Table 7B Red deer culling model for the CSDMG eastern sub-area plus Invercauld (part)
Year Population Stag Hind Calf Total Density
(km-2)*
Density
(km-2)**
2005 Spring count 772 1331 493 2596   5.4
  Recruitment (Yearlings) 246 247        
  Summer (after calving) 1018 1578 631 3227 5.2  
  Cull (=recruitment) 246 247 98 591    
2006 Spring estimate 772 1331 493 2596   5.4
  Ongoing maintenance cull = recruitment 246 247 98      

* Summer densities based on the entire available range of 62,328 ha.
** Spring densities are computed based on a winter range of 47,814 ha.

Assumes 0.4 of hinds in summer calving; 0.37 of calves recruited into adult population in spring..
Figures for the numbers of calves in the cull and in summer are crude estimates based on numbers of hinds. Calves should be culled in relation to milk hinds in the cull.

Notes for the application of these models:

Although this model can be modified annually following future spring counts, a strong commitment should be given to achieving the proposed reduction.

Consideration might be given to further modifying the stag to hind ratio in favour of stags, so as to provide more mature stags while maintaining a low total population size. A bias in favour of stags will assist the achievement of sporting targets while maintaining a population density consistent with other estate objectives.

It is extremely unlikely that red deer numbers will be reduced to levels below which they can make a speedy recovery, with a reduction in culling effort, should this be considered necessary at any stage.

Local (estate-based) culls aimed at maintaining the CSDMG eastern sub-area density of 3.4 deer km-2 (Table 7A), are proposed in Table 8. The division of the total cull requirement between estates has been based on the proportion of low ground as representing the approximate carrying capacity of the habitat. These proposed culls are indicative only and estates should share information on culling progress during culling seasons aimed at achieving the sub-area total.

 
Table 8 Indicative proposed red deer culls for estates in the eastern sub-area
Estate % low ground Stag Hind Calf Total
Total** 100 93 94 38

225

** Total cull figures are taken from Table 7A (Maintenance cull)

Website: Ardchattan Solutions Ltd